Marine Biodiversity » Marine Sponges » Calcarea » Calcarea Classification » Subclass Calcaronea

Marine SpongesCalcarea Introduction
Calcarea ClassificationSubclass CalcineaSubclass Calcaronea

Order Leucosoleniida
Family Leucosoleniidae
Family Sycettidae
Family Heteropiidae
Family Grantiidae
Family Amphoriscidae
Family Lepidoleuconidae
Family Staurorrhaphidae
Order Lithonida

Subclass Calcaronea
Order Leucosoleniida
Family Leucosoleniidae MINCHIN, 1898

taken from 'Sponguide' by J.N.A. Hooper (Queensland Museum, Brisbane)

Definition: Asconoid, erect growth forms, with long, individual, clustered, oscular tubes arising from stolon-like system of basal tubes; tubes may have diverticular and often arborescent; monaxon spicules always present; triradiates, if present, typically bilateral, sagittal, inequiangular in form (where two of the rays form a pair, while the third differs in some way), and with the crystalline optic axis never vertical but always inclined to the facial plane of the rays; choanocytes with flagellum arising directly from the pear-shaped nucleus, situated at or near the apex of the cell; choanocytes line central cavity (spongocoele) of the individual tubes; larvae are amphiblastula

Scope: About 90 generic names are included in this family, but only three of these are now.

Reviews: Bidder (1898), Minchin (1909), Dendy & Row (1913), Hartman (1958), Burton (1963).

Genera:
Ascute Dendy & Row, 1913 (type species: Leucosolenia uteoides Dendy, 1893) - compound, tubular, sessile, surface even, minutely hispid, striated; oscules apical; firm texture; ectosomal skeleton has large diacts (oxeotes), of one or two sizes, echinating surface and/or lying longitudinally within the ectosome; choanosomal skeleton composed of sagittal quadriradiates (Burton, 1963).
Ascyssa Haeckel, 1872 (type species: Ascyssa acufera Haeckel, 1872) - tubular, colonial; smooth surface; apical oscules; skeleton a tangential layer of irregularly arranged oxeote and microxeote spicules (Burton, 1963).
Leucosolenia Bowerbank, 1861 (type species: Spongia botryoides Ellis & Solander, 1786) (syn. Nardosa Wright, 1868; Olynthium Haeckel, 1870; Olynthus Haeckel, 1870; Prosycum Haeckel, 1870; Leucaria Haeckel, 1870; Leucelia Haeckel, 1870; Leuceria Haeckel, 1870; Leucilia Haeckel, 1870; Leuciria Haeckel, 1870; Leucoria Haeckel, 1870; Tarroma Haeckel, 1870; Tarrus Haeckel, 1870; Nardopsis Haeckel, 1870; Clystolynthus Haeckel, 1870; Sycorrhiza Haeckel, 1870; Auloplegma Haeckel, 1870; Thecometra Haeckel, 1870; Ascettaga Haeckel, 1872; Ascettopa Haeckel, 1872; Ascettusa Haeckel, 1872; Ascometra Haeckel, 1872; Nardorus Haeckel, 1872; Solenula Haeckel, 1872; Tarropsis Haeckel, 1872; Ascilla Haeckel, 1872; Ascillaga Haeckel, 1872; Ascillopa Haeckel, 1872; Ascaltaga Haeckel, 1872; Ascaltopa Haeckel, 1872; Ascuris Haeckel, 1872; Ascortaga Haeckel, 1872; Ascortopa Haeckel, 1872; Ascortusa Haeckel, 1872; Asculmis Haeckel, 1872; Ascandraga Haeckel, 1872; Ascandropa Haeckel, 1872; Olynthella Haeckel, 1872; Nardoma Haeckel, 1872; Solenidium Haeckel, 1872; Olynthaltus Haeckel, 1872; Olynthandrus Haeckel, 1872; Olynthettus Haeckel, 1872; Olynthortus Haeckel, 1872; Olynthillus Haeckel, 1872; Olynthulmus Haeckel, 1872; Olynthandrium Haeckel, 1872; Olynthandra Haeckel, 1872; Olynthelletta Haeckel, 1872; Clistolynthus Haeckel, 1872; Clistolynthaltis Haeckel, 1872; Clistolynthandra Haeckel, 1872; Clistolynthetta Haeckel, 1872; Clistolynthilla Haeckel, 1872; Soleniscetta Haeckel, 1872; Soleniscilla Haeckel, 1872; Soleniscyssa Haeckel, 1872; Solenicaltis Haeckel, 1872; Solenicortis Haeckel, 1872; Soleniculmis Haeckel, 1872; Soleniscandra Haeckel, 1872; Solenidandra Haeckel, 1872; Solenulandra Haeckel, 1872; Solenuletta Haeckel, 1872; Nardoraltis Haeckel, 1872; Nardorandra Haeckel, 1872; Nardoretta Haeckel, 1872; Nardorilla Haeckel, 1872; Nardorortis Haeckel, 1872; Nardomandrum Haeckel, 1872; Nardopsandra Haeckel, 1872; Nardopsetta Haeckel, 1872; Nardopsortis Haeckel, 1872; Tarraltis Haeckel, 1872; Tarrandra Haeckel, 1872; Tarretta Haeckel, 1872; Tarrilla Haeckel, 1872; Tarromandra Haeckel, 1872; Tarropsandra Haeckel, 1872; Tarropsetta Haeckel, 1872; Auloplegmaltis Haeckel, 1872; Auloplegmetta Haeckel, 1872; Auloplegmilla Haeckel, 1872; Auloplegmandra Haeckel, 1872; Auloplegmortis Haeckel, 1872; Ascaltometra Haeckel, 1872; Ascandrometra Haeckel, 1872; Ascettometra Haeckel, 1872; Homandra Lendenfeld, 1891; Nardoris Delage, 1899; Aulorhiza Dendy & Row, 1913) - encrusting network of more or less erect asconoid tubes which may branch, or throw out lateral diverticula, but seldom anastomose except occasionally at base of main tubes; surface minutely hispid; texture soft; skeleton of sagittal triradiates and sagittal quadriradiates, with apical rays projecting into cloacal cavity, and oxeas with lanceolate distal ends (Burton, 1963).